Recent Publications


  1. Balasubramaniam KN, Marty PR, Arlet M, Beisner BA, Kaburu SSK, Bliss-Moreau E, Kodandaramaiah U, McCowan B. (2020). Impact of anthropogenic factors on affiliative behaviors among bonnet macaques American Journal of Physical Anthropology,
  2. Vandeleest, J.J., Winkler, S.L., Beisner, B.A., Hannibal, D.L., Atwill, E.R., McCowan, B. (2020). Sex differences in the impact of social status on hair cortisol concentrations in rhesus monkeys (Macaca mulatta). American Journal of Primatology.
  3. Beisner B, Posfai M, Braun N, D’Souza RM, McCowan B. (2020). Individual attributes predict multiplex centrality in rhesus macaques, PeerJ.


  1. Beisner BA, Wooddell LJ, Hannibal DL, Nathman A, McCowan B. (2019). High rates of aggression do not predict rates of trauma in captive groups of macaques.Applied Animal Behavior Science,
  2. Posfai M, Braun N, Beisner B, McCowan B, D’Souza RM (2019) Consensus ranking for multi-objective interventions in multiplex networks. New Journal of Physics,
  3. Miller E, Johnson T, Omondi G, Atwill E, Isbell L, McCowan B, and VanderWaal K (2019). Assessing transmission of antimicrobial resistant Escherichia coli in wild giraffe contact networks. Applied and Environmental Microbiology,
  4. Marty PR, Beisner B, Kaburu SSK, Balasubramaniam K, Bliss-Moreau E, Ruppert N, Sah SAM, IsmailA, ArletME, AtwillER, McCowan, B (2019). Time constraints imposed by anthropogenic environments alter social behaviour in long-tailed macaques. Animal Behaviour, 150:157-165.
  5. Balasubramaniam K, Beisner B, Hubbard J, Vandeleest J, Atwill ER, McCowan B (2019). Affiliation and disease risk: social networks mediate gut microbial transmission among rhesus macaques. Animal Behaviour,
  6. Wooddell LJ, Beisner B, Hannibal D, Nathman A, McCowan B (2019). Increased produce enrichment reduces trauma in socially-housed captive rhesus macaques (Macaca mulatta). American Journal of Primatology.
  7. Kaburu S, Beisner B, Balasubramaniam K, Marty P, Bliss-Moreau E, Atwill ER, Arlet M, McCowan B (2019). Interactions with humans impose time constraints on urban-dwelling rhesus macaques (Macaca mulatta). Behavior,
  8. Marty PR, Balasubramaniam KN, Kaburu SSK, Hubbard J, Beisner B, Bliss-Moreau E, Ruppert N, Arlet ME, Sah SAM, Ahmad Ismail, Mohan L, Rattan SK, Kodandaramaiah U, McCowan B. (2019). Individuals in urban dwelling primate species face unequal benefits associated with living in an anthropogenic environment. Primates,


  1. McCowan B, Beisner BA, Hannibal DL (2018) Social management of laboratory rhesus macaques housed in large groups using a network approach: a review. Behavioral Processes
  2. Balasubramaniam KN, Beisner BA, Berman CM, De Marco A, Duboscq J, Koirala S, Majolo B, MacIntosh AJ, McFarland R, Molesti S, Ogawa H, Petit O, Schino G, Sosa S, Sueur C, Thierry B, de Waal FBM, McCowan B (2018) The influence of phylogeny social style and sociodemographic factors on variation in macaque social networks. American Journal of Primatology
  3. Balasubramaniam K, Beisner B, Guan J, Vandeleest J, Fushing H, Atwill E, McCowan B. (2018) Social network community structure and the contact-mediated sharing of commensal E. coli among captive rhesus macaques (Macaca mulatta) PeerJ 6:e4271,
  4. Cassidy L, Hannibal D, Vandeleest J, Semple S, Barnard A, Chun K, Winkler S, McCowan B (2018). Intermittent pair-housing and pair relationship qualities modulate HPA activity in adult female rhesus macaques. Applied Animal Behaviour Science,
  5. Fushing H, Liu S, McCowan B (2018). From patterned response dependency to structured covariate dependency: entropy based categorical-pattern-matching. PLoS One 13: e0198253.
  6. Gottlieb DL, Capitanio JP, McCowan B (2018). Personality, environmental stressors, and diarrhea in rhesus macaques: an interactionist perspective. American Journal of Primatology,
  7. Kaburu S, Beisner B, Balasubramaniam K, Marty P, Bliss-Moreau E, McCowan B (2018). Rates of human-macaque interactions affect grooming behavior among urban-dwelling rhesus macaques (Macaca mulatta). American Journal of Physical Anthropology, 168: 92-103,
  8. Vandeleest JJ, Capitanio JP, Hamel A, Meyer J, Novak M, Mendoza SP, McCowan B (2018). Social stability influences the association between adrenal responsiveness and hair cortisol concentrations in rhesus macaques, Psychoneuroendocrinology,
  9. Linden J, McCowan B, Capitanio J, Isbell L (2018). Male-inflicted wounds have opposite effects on hair cortisol for captive male and female rhesus macaques (Macaca mulatta) following new group formation. Primates,
  10. Linden J, Capitanio JP, McCowan B, Isbell LA (2018). Coping style and cortisol levels in infancy predict hair cortisol following new group formation in captive rhesus macaques (Macaca mulatta). American Journal of Primatology,


  1. Hannibal DL, Bliss-Moreau E, Vandeleest J, McCowan B, Capitanio J (2017) Laboratory rhesus macaque social housing and social changes: Implications for research. American Journal of Primatology;
  2. Capitanio JP, Blozis SA, Snarr J, Steward A, McCowan B (2017) Do “birds of a feather flock together” or do “opposites attract”? Behavioral responses and temperament predict success in pairings of rhesus monkeys in a laboratory setting. American Journal of Primatology;
  3. Theil JH, Beisner BA, HillAE, McCowanB (2017) Effects of human management events on conspecific aggression in captive rhesus macaques (Macaca mulatta). Journal of the American Association for Laboratory Animal Science 56 109.
  4. Seil SK, Hannibal DL, Beisner BA, McCowan B (2017) Predictors of insubordinate aggression among captive female rhesus macaques. American Journal of Physical Anthropology